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Ashitaka finds himself afflicted by a mysterious illness. He sets off west in search of it and meets San, a young girl who lives with the wolves.
When two girls move to the country to be near their ailing mother, they have adventures with the wondrous forest spirits who live nearby.
With intense acting and a haunting atmosphere, the film is an emotional rollercoaster that confronts the viewer with themes of shame, guilt and forgiveness.